The other example is the thumb of the giant
panda, another favorite example of Gould's.
The giant panda has a thumb that it uses to strip
leaves from bamboo stalks in order to make them
easier to eat.
It can't move much, but can squeeze against the
fingers (and true thumb) to hold bamboo.
http://www.athro.com/evo/pthumb.html
However, the "thumb" is not really a digit
but rather an enlarged radial sesamoid,
a bone that grows in joints.
Sounds like a pretty good adaptation, right? Giant
pandas evolve an enlarged radial sesamoid in order
to better eat bamboo.
http://www.athro.com/evo/pthumb.html
It turns out that grizzly bears do not have this enlarged
radial sesamoid. It also turns out that giant pandas
have this enlarged radial sesamoid in their hind feet
as well.
What does this imply?